Principal / Senior Software Engineer
Flagship Pioneering, Inc., Cambridge, MA USA +1
Flagship Pioneering, Inc., Cambridge, MA USA +1
We are seeking an experienced, creative, and talented Principal / Senior Software Engineer. The ideal candidate will have a strong background in software and product development, with the ability to lead projects and communicate effectively across teams and with ML researchers.
Successful candidates will be able to thrive in a fast-paced, action-oriented, team environment and display a clear passion for AI innovation and impact. The role offers a unique opportunity to help drive the strategy of an early stage AI-focused company. The candidate must be an entrepreneurial, highly motivated, and purpose-driven individual. The successful candidate will have a track record of building consumer applications or large internal applications. They must exhibit strong collaboration and communication skills. They should be ready to lead application development in a very dynamic and innovative environment.
The Company
Iris Labs, Inc. is a privately held, early-stage company pioneering the use of artificial intelligence to transform how people navigate life’s challenges and wellbeing. We are creating a platform that empowers people to build a life they are proud of and fulfilled by. We bring together psychology, product, design, engineering, and machine learning to build digital experiences that are engaging, trustworthy, and genuinely helpful.
Iris Labs is backed by Flagship Pioneering, an innovation enterprise that conceives, creates, resources, and builds companies that invent breakthrough technologies that transform the world. Flagship has created over 100 groundbreaking companies since 2000, including Moderna.
Flagship Pioneering is a bioplatform innovation company that invents and builds platform companies, each with the potential for multiple products that transform human health or sustainability. Since its launch in 2000, Flagship has originated and fostered more than 100 scientific ventures, resulting in more than $90 billion in aggregate value. Many of the companies Flagship has founded have addressed humanity’s most urgent challenges: vaccinating billions of people against COVID-19, curing intractable diseases, improving human health, preempting illness, and feeding the world by improving the resiliency and sustainability of agriculture.
Flagship has been recognized twice on FORTUNE’s “Change the World” list, an annual ranking of companies that have made a positive social and environmental impact through activities that are part of their core business strategies, and has been twice named to Fast Company’s annual list of the World’s Most Innovative Companies. Learn more about Flagship at www.flagshippioneering.com.
At Flagship, we recognize there is no perfect candidate. If you have some of the experience listed above but not all, please apply anyway. Experience comes in many forms, skills are transferable, and passion goes a long way. We are dedicated to building diverse and inclusive teams and look forward to learning more about your unique background.
Recruitment & Staffing Agencies: Flagship Pioneering and its affiliated Flagship Lab companies (collectively, “FSP”) do not accept unsolicited resumes from any source other than candidates. The submission of unsolicited resumes by recruitment or staffing agencies to FSP or its employees is strictly prohibited unless contacted directly by Flagship Pioneering’s internal Talent Acquisition team. Any resume submitted by an agency in the absence of a signed agreement will automatically become the property of FSP, and FSP will not owe any referral or other fees with respect thereto.
The salary range for this role is $108,000 - $209,000. Compensation for the role will depend on a number of factors, including a candidate’s qualifications, skills, competencies, and experience. Iris Labs, Inc. currently offers healthcare coverage, annual incentive program, retirement benefits and a broad range of other benefits. Compensation and benefits information is based on Iris Labs, Inc.'s good faith estimate as of the date of publication and may be modified in the future.
Flagship Pioneering and our ecosystem companies are committed to equal employment opportunity regardless of race, color, ancestry, religion, sex, national origin, sexual orientation, age, citizenship, marital status, disability, gender identity or Veteran status.
LightfieldHQ: San Francisco
$220K–$300K / yr
Posted Oct 13, 2026
LightfieldHQ: San Francisco
$220K–$300K / yr
Posted Oct 7, 2026
BrandlightTLV Office
Pay not posted
Posted 13 days ago
BrandlightTLV Office
Pay not posted
Posted 12 days ago
Lila SciencesCambridge, MA USA
Source: the Flagship Pioneering, Inc. Greenhouse job board. We last checked this job 7 h ago. We first saw it 8 days ago. A job on a job board does not always mean that the team interviews people now. We found this job board in a public list. We did not yet compare it with the careers page of the company.
Privacy Notice for Applicants: When you apply for a role at Flagship Pioneering or one of its portfolio companies, we collect and use personal information you provide (such as your name, contact details, work history, and application materials) to evaluate your application, communicate with you, and comply with legal obligations. Your application data is processed through Greenhouse, our applicant tracking system, and may also be reviewed using AI-assisted screening tools.
We do not sell your personal information. California residents have rights under the CCPA/CPRA including to know, delete, and opt out of the sharing of their personal information. If you are located in the EU or UK, we process your data under GDPR and you have rights to access, rectify, and erase your data. To exercise your rights or for questions, contact privacy@flagshippioneering.com.
$400K–$500K / yr
Posted Apr 8, 2026
Raspberry AIUnited States
Pay not posted
Posted Feb 3, 2026
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This base pay is below the median for principal software engineering jobs in other US places.
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